Notholaena jacalensis
Pray's Cloakfern
Carl RothfelsIntroduction
I know of only two locations for this unusual species. It was first described from chalky limestone roadside rockcuts outside of the town of Jacala, Hidalgo (Pray, 1967), where it is still locally common today (at least as recently as August 2009). It is also found (and is also locally common) in the gypsum outcrops of Municipio San Nicolas Tolentino, San Luis Potosí, which it shares with the extremely local endemic Pellaea ribae (known from no other location). At the latter site, N. jacalensis plants can be quite large, approaching N. copelandii in size.
Characteristics
Notholaena jacalensis most closely resembles a small version of N. copelandii, and thus its phylogenetic position with N. bryopoda and N. neglecta is rather surprising. Interestingly, in the original description, Pray (1967) suggests that N. jacalensis shares a resemblance with N. greggii (and thus with N. bryopoda by extension), an observation worth further attention in light of our molecular data. From Notholaena copelandii it differs in its narrower leaf, more-gradual tapering of the leaf tip, widely spaced segments, and purported by farina colour (with N. jacalensis having light yellow farina, versus the white farina of N. copelandii). However, in the field, the farina colour of N. jacalensis seemed to vary based on exposure and age, with some individuals distinctly yellowish, and others nearly white. Notholaena jacalensis also differs from N. copelandii in its rhizome scales, which are smooth, and resemble those of N. candida. With Notholaena bryopoda and greggii it shares the widely spaced segments, and the slight narrowing of the bases of the ultimate segments (Pray 1967).
Discussion of Phylogenetic Relationships
Single-locus chloroplast data (Rothfels unpublished) place N. jacalensis in a polytomy with N. bryopoda and N. neglecta, well within core Notholaena II.
References
Mickel, J. T., and A. R. Smith. 2004. The Pteridophytes of Mexico. The New York Botanical Garden Press, New York.
Pray, T. R. 1967. A new species of Notholaena from Mexico. American Fern Journal 57:101--103.
Title Illustrations

| Scientific Name | Notholaena jacalensis | 
|---|---|
| Location | San Luis Potosí, Mexico | 
| Specimen Condition | Live Specimen | 
| Identified By | C.J.Rothfels | 
| Image Use |  This media file is licensed under the Creative Commons Attribution-NonCommercial License - Version 3.0. | 
| Copyright | © 2010 Carl Rothfels   | 
About This Page
	
        Carl Rothfels
    
	   
    
    
	 
 
    
	Duke University, Durham, North Carolina, USA
				Correspondence regarding this page should be directed to Carl Rothfels at 
            
Page copyright © 2010 Carl Rothfels
    
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    Notholaena jacalensis .  Pray's Cloakfern. 
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    Page: Tree of Life 
    Notholaena jacalensis .  Pray's Cloakfern. 
    Authored by 
    Carl Rothfels.         
	The TEXT of this page is licensed under the 
    
    Creative Commons Attribution-NonCommercial License - Version 3.0. Note that images and other media 
    featured on this page are each governed by their own license, and they may or may not be available 
    for reuse. Click on an image or a media link to access the media data window, which provides the 
    relevant licensing information. For the general terms and conditions of ToL material reuse and 
    redistribution, please see the Tree of Life Copyright 
    Policies.
- First online 07 January 2010
- Content changed 07 January 2010
Citing this page:
Rothfels, Carl. 2010. Notholaena jacalensis . Pray's Cloakfern. Version 07 January 2010 (under construction). http://tolweb.org/Notholaena_jacalensis/133609/2010.01.07 in The Tree of Life Web Project, http://tolweb.org/

 
			
	    
		 
        	
	
		 
			
	
		

 
     
 
	
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