Josiini
James S. Miller and Andrew V. Z. Brower
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close boxIntroduction
A group of neotropical day-flying moths with black and yellow aposematic color patterns.Characteristics
Josiini possess a unique metathoracic tympanal organ that separates them from other moths. In addition, rami of the antennae are joined to the antennal shaft with hingelike joints, fore- and hindwing veins M3 and CuA1 are stalked, and forewings lack stridulatory organs (Miller 2009).
Larvae have a glassy surface to the head capsule and go through four instars. Pupae on the inderside of leaves are surrounded by a loose net of silk.
A diagram of josiine thoracic structures. Note the kettle drum-shaped metathoracic tympanum.
Caribojosia youngi. © 2009 James S. Miller
Discussion of Phylogenetic Relationships
The hypothesis of relationships among josiine genera is based on a cladistic analysis of larval and adult morphological characters (Miller 2009).References
Miller JS. 1996. Phylogeny of the Neotropical moth tribe Josiini (Notodontidae: Dioptinae): a hidden case of Müllerian mimicry. Zool. J. Linn. Soc. 118, 1-45.
Miller, JS. 2009. Generic revision of the Dioptinae (Lepidoptera: Noctuoidea: Notodontidae). Bulletin of the American Museum of Natural History 321, 1-971 + 48 plates.
Miller JS, Brower AVZ. & Desalle R. 1997. Phylogeny of the neotropical moth tribe Josiini (Notodontidae: Dioptinae): comparing and combining evidence from DNA sequences and morphology. Biol. J. Linn. Soc. 60, 297-316.
Title Illustrations
| Scientific Name | Lyces flavissima |
|---|---|
| Location | Venezuela |
| Reference | Miller JS. 2009. Generic revision of the Dioptinae (Lepidoptera: Noctuoidea: Notodontidae). Bulletin of the American Museum of Natural History 321, 1-971 + 48 plates. (Plate 39) |
| Creator | L. D. Otero |
| Specimen Condition | Live Specimen |
| Identified By | James S. Miller |
| Life Cycle Stage | larva |
| View | dorsal-lateral |
| Image Use |
 This media file is licensed under the Creative Commons Attribution License - Version 3.0.
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| Copyright |
© 2009
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| Scientific Name | Josia sp. |
|---|---|
| Location | Ecuador |
| Comments | Note the net of silk strands surrounding the pupa. |
| Reference | Miller JS. 2009. Generic revision of the Dioptinae (Lepidoptera: Noctuoidea: Notodontidae). Bulletin of the American Museum of Natural History 321, 1-971 + 48 plates. (Plate 39) |
| Creator | J. E. Rawlins |
| Specimen Condition | Live Specimen |
| Identified By | James S. Miller |
| Life Cycle Stage | pupa |
| View | lateral |
| Image Use |
This media file is licensed under the Creative Commons Attribution License - Version 3.0.
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| Copyright |
© 2009
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| Scientific Name | Scea superba |
|---|---|
| Location | Ecuador: Napo, Yanayacu Biological Station, 5 km w Cosanga 00°35.9'S, 77°53.4' W 2163 m. |
| Reference | Miller JS. 2009. Generic revision of the Dioptinae (Lepidoptera: Noctuoidea: Notodontidae). Bulletin of the American Museum of Natural History 321, 1-971 + 48 plates. (Plate 34) |
| Specimen Condition | Live Specimen |
| Identified By | James S. Miller |
| Sex | Male |
| Life Cycle Stage | adult |
| View | dorsal |
| Collection | AMNH |
| Collector | Harold Greeney et al. |
| Image Use |
This media file is licensed under the Creative Commons Attribution License - Version 3.0.
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| Copyright |
© 2009
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About This Page
Middle Tennessee State University, Murfreesboro, Tennessee, USA
Correspondence regarding this page should be directed to James S. Miller at and Andrew V. Z. Brower at
Page copyright © 2010 and
Page: Tree of Life
Josiini .
Authored by
James S. Miller and Andrew V. Z. Brower.
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- First online 12 August 2009
- Content changed 10 January 2010
Citing this page:
Miller, James S. and Andrew V. Z. Brower. 2010. Josiini . Version 10 January 2010 (under construction). http://tolweb.org/Josiini/138572/2010.01.10 in The Tree of Life Web Project, http://tolweb.org/
